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His father Fe’ao captained Tonga in the 1990s when several uncles also represented the national side. His cousin Toby stars for Wales, while Vunipola and his brother Mako are standout players for England.
But that hasn’t always made things easy, particularly during his early years.
Born in Sydney, raised in Wales and then England when the family relocated to the UK – Fe’ao had signed to play for Pontypool – the imposing number eight Billy describes his itinerant upbringing as “frantic,” adding, “we were never settled anywhere until we came to England.” Britain proved something of a culture shock.
“When we started setting up roots here, it was tough,” says Vunipola. “I remember our first few months in Wales we all just slept in the living room because we were used to being around each other. We just had duvets all over the floor.”
It’s an endearing image and one that’s difficult to picture today.
Vunipola (right) celebrates England's Six Nations victory over Scotland in 2016, when the team also won the grand slam.
Vunipola tips the scales at 130 kg. He’s a bulldozer with the ball in hand, but has honed his game in recent years to develop defensive nous and subtle handling skills.
Brother Mako, meanwhile, two years Billy’s senior, is 9 kg lighter but equally adept in open play, eschewing the logic that prop forwards are supposed to focus solely on scrummaging.
Both will feature prominently in England’s upcoming Six Nation campaign, which gets underway in Dublin against Ireland on February 2, with Vunipola having recently returned from a long injury layoff that saw him suffer three arm breaks.
“My name’s on the sheet but I want to be back back,” he says. “I want to be playing and I want to be playing well.
“I’m lucky that I only had to deal with broken arms, you know. I can still walk and I can still play rugby. It just took time out. It wasn’t fun but I’m happy, I’m playing rugby and hopefully I can carry on.
“The Six Nations is probably the premier tournament outside the World Cup in terms of rugby, I would say, and in terms of viewership. The rugby that’s been played, the unpredictability of the whole tournament, just as a fan it’s exciting. As a player it’s even more exciting.”
After a disappointing fifth-place finish in last year’s tournament – for which Vunipola was absent through injury – Eddie Jones’ England side will be hoping for a much-improved Six Nations campaign with a World Cup on the horizon, and Vunipola’s return should give confidence in that regard.
Since making his international debut in 2013, he’s been a leading light in an England team that’s endured unprecedented high and record lows, as well as helping his club side Saracens amass silverware on domestic and European fronts.
A rugby career, he says, always seemed inevitable.
“I remember growing up and my dad trying to push us in the direction of being doctors or lawyers or anything he thought was a good job,” says Vunipola. “I’m not saying he was right or wrong but me and my brother always wanted to play rugby. We got our wish.”
The seed was sown when the brothers started to wrestle with a rugby ball in their living room in Wales or played mini games with their cousins outdoors.
“Mako would be with Toby, who’s now at Bath, and I would be with Toby’s brother,” recalls Vunipola. “Every single time we played it was always me and Mako that had a fight.
“I was always the youngest of the four of us. They’d both hate it if I actually got a successful tackle on them. He’d usually start [the fights] and finish them, but I knew that I was getting under his skin when I’d tackle him or score a try.”
Billy and Mako celebrate with their family after winning the European Champions Cup in 2017.
Stu Forster/Getty Images Europe/Getty Images
That rivalry has mellowed now, with Vunipola even admitting that his brother is the superior player: “He’s probably just a smaller, better, stronger version of me,” he says.
Humility is something Vunipola has acquired over time. He says that his rise to the top of the game – via a scholarship at prestigious English public school Harrow and the Wasps academy – gave him undue self-confidence.
“If you keep hearing your name I guess subconsciously you just think you’re the man and that’s what I did. I thought I was more important than I actually was.
“I love seeing some of the young boys here at the club. You go in 50% and they go 100% and you get angry at them, but they do what’s right. I’ve actually learned a lot from the younger guys coming in.”
New Zealand beat Australia 34-17 to win its second straight Rugby World Cup at Twickenham Stadium, London in October 2015. Four years on, the focus will shift to Japan, where 12 stadiums throughout the country will host the tournament from September 20 to November 2.
Stu Forster/Getty Images
What: International Stadium Yokohama Capacity: 72,327 Where: Yokohama City, Kanagawa Prefecture Matches: New Zealand vs South Africa; Ireland vs Scotland; England vs France; Japan vs Scotland; Semifinals 1 & 2; Final
Rugby World Cup 2019
What: Tokyo Stadium Capacity: 49,970 Where: Tokyo Metropolitan Matches: Japan vs Romania; France vs Argentina; Australia vs Wales; England v Argentina; New Zealand vs Namibia; Quarterfinals 2 & 4; Bronze final
Rugby World Cup 2019
What: City of Toyota Stadium Capacity: 45,000 Where: Toyota, Aichi Prefecture Matches: Wales vs Georgia; South Africa vs Namibia; Japan vs Samoa; New Zealand vs Italy
Rugby World Cup 2019
What: Shizuoka Stadium Ecopa Capacity: 50,889 Where: Shizuoka Prefecture Matches: Japan vs Ireland; South Africa vs Italy; Scotland vs Romania; Australia vs Georgia
Rugby World Cup 2019
What: Hanazono Rugby Stadium Capacity: 32,228 Where: Higashiosaka City, Osaka Prefecture Matches: Italy vs Namibia; Argentina vs Tonga; Georgia vs Fiji; USA vs Tonga
Rugby World Cup 2019
What: Kumamoto Stadium Capacity: 32,228 Where: Kumamoto City, Kumamoto Prefecture Matches: France vs Tonga; Wales vs Uruguay
Rugby World Cup 2019
What: Oita Stadium Capacity: 40,000 Where: Oita Prefecture Matches: New Zealand vs Canada; Australia vs Uruguay; Wales vs Fiji; Quarterfinals 1 & 3
Rugby World Cup 2019
What: Kumagaya Rugby Stadium Capacity: 25,600 Where: Kumagaya City, Saitama Prefecture Matches: Russia vs Samoa; Georgia vs Uruguay; Argentina vs USA
MARTIN BUREAU/AFP/AFP/Getty Images
What: Kamaishi Recovery Memorial Stadium Capacity: 16,334 Where: Kamaishi City, Iwate Prefecture Matches: Fiji vs Uruguay; Namibia vs Canada
QUENTIN TYBERGHIEN/AFP/AFP/Getty Images
What: Sapporo Dome Capacity: 41,410 Where: Sapporo City Matches: Australia vs Fiji; England vs Tonga
Rugby World Cup 2019
What: Kobe Misaki Stadium Capacity: 30,132 Where: Kobe City Matches: England vs USA; Scotland vs Samoa; Ireland vs Russia; South Africa vs Canada
Rugby World Cup 2019
What: Fukuoka Hakatanomori Stadium Capacity: 21,562 Where: Fukuoka City, Fukuoka Prefecture Matches: Italy vs Canada; France vs USA; Ireland vs Samoa
Rugby World Cup 2019
Rugby World Cup 2019: The host venues in Japan
While his initial focus is having an injury-free run of games through the Six Nations, Vunipola admits that his thoughts can’t help but gravitate towards the World Cup in Japan later this year.
He was part of the England team that crashed out at the pool stages in 2015 – the first host nation ever to do so – and will be eager to right some wrongs in Japan.
“This is what you kind of prepare your whole life for,” he says, “I guess I didn’t understand that in 2015.”
“We’ve got a Six Nations coming up, but, you know, it’s something you can’t be afraid to talk about. It’s the World Cup, it’s massively exciting. It’s going to be a massive year for everyone.”
New Zealand became the first side to retain the Rugby World Cup after defeating Australia 34-17 in the final at Twickenham Stadium.
Mike Hewitt/Getty Images Europe/Getty Images
New Zealand, 2011 —
The All Blacks ended a 24-year drought when they lifted the Web Ellis trophy on home soil in 2011. Here, Ma'a Nonu greets raucous crowds during the victory parade in Wellington.
Phil Walter/Getty Images AsiaPac/Getty Images
France, 2007 —
South Africa claimed its second title in France in 2007, defeating England 6-15 at the Stade de France in Paris.
Gallo Images/Getty Images Europe/Getty Images
Australia, 2003 —
England became the first -- and to this day the only -- northern hemisphere side to win the World Cup in 2003 by defeating Australia 20-17 in Sydney. Fly-half Jonny Wilkinson struck the winning drop goal in extra-time.
DAMIEN MEYER/AFP/AFP/Getty Images
Wales, 1999 —
Australia's second World Cup victory in 1999 came eight years after its first. A 35-12 victory over France saw John Eales' side lift the trophy in Cardiff.
OLIVIER MORIN/AFP/AFP/Getty Images
South Africa, 1995 —
It's one of rugby's most iconic images -- South African President Nelson Mandela presenting the World Cup to Springbok captain Francois Pienaar in 1995. The victory helped unite the nation shortly after the end of apartheid.
JEAN-PIERRE MULLER/AFP/AFP/Getty Images
England, 1991 —
Australia first tasted World Cup victory in 1991 after narrowly defeating England 12-6 in the final.
Russell Cheyne/Getty Images Europe/Getty Images
New Zealand, 1987 —
The All Blacks won the inaugural Rugby World Cup on home soil in 1987. Captain David Kirk kisses the trophy following victory over France in Auckland.
GEORGES GOBET/AFP/AFP/Getty Images
Rugby World Cup: The teams to lift the Webb Ellis trophy
Japan 2019
England’s opening match of the tournament – against Tonga – will divide the Vunipola household. Fe’ao won 32 caps for the South Pacific island and today works for the Tonga Rugby Union.
“I guess we’re his sons and he’ll want us to do well but you know deep down inside he wants Tonga to secretly get one over England,” says Vunipola.
“I know my mom will 100% be behind us and the rest of my family but my Dad will be a bit torn.”
Born in Australia, raised in the UK and crowned world champion in Japan? Vunipola’s story would enter another thrilling chapter if England were to defy the odds and lift the World Cup.
“The thing is I’m one of millions of people that think about winning in Japan and I guess I’m one of the millions that aren’t scared to talk about it,” he says.
“All these cliché things that you can sit here and say in an interview is correct for right now. It’s a dream.”